Wagner, Warren L., Derral R. Herbst and S. H. Sohmer. 1999. Manual of the Flowering Plants of Hawai'i (Bernice Pauahi Bishop Museum Special Publication, 2 Vol. Set). Honolulu, HI
Flora of Hawaii:
Distribution In the Hawaiian Islands, endemic to Lana`i.
Habit Shrubs, closely grayish pubescent with stellate and simple glandular hairs.
Leaves Leaf blades deeply cordate-ovate, 7-12 cm long, margins crenate-dentate, apex acute to acuminate, petioles about as long as blades, sometimes pilose with simple hairs. Flowers
Flowers Flowers solitary or paired in the leaf axils, exceeded by the leaves, pedicels 1.5-4 cm long, articulate with the calyx sometimes pilose with simple hairs and stellate and glandular pubescent; calyx cup-shaped to suburceolate, 1-1.5 cm long, lobes dentate, 1-3 mm long, pubescent within; petals included in calyx, greenish with brighter green, inflexed margins, lower surface reddish, 0.5-1 cm long, up to 9 mm wide; staminal column exserted, whitish to yellowish, 2-2.8 cm long, antheriferous in a terminal cluster; style branches coral-colored; stigmas green, scarcely expanded.
Fruit Schizocarp cylindrical, 8-10 mm long, 10-13 mm in diameter, pilose, mericarps 8-9, 8-10 mm long, apex acute.
Seeds Seeds asymmetrically reniform, up to 2.8 mm long, matted white or brown pubescent.