(nat) Pualele
Annual herbs 1-20 dm tall, subglabrous, sometimes with a few glandular hairs in the inflorescence. Leaves lanceolate, cordate- lanceolate to spatulate, 6-30 cm long, 1-15 cm wide, usually lyrate-pinnatifid, sometimes merely toothed, sessile and auriculate, the auricles acute at base. Involucre 9-13 mm high in fruit. Achenes transversely rugulose and 6-10-ribbed, 2.5-3 mm long, ca. 3-4 times as long as wide. [2n = 16, 18, 32, 36.] Native to Europe, now a cosmopolitan weed; in Hawaii naturalized in a variety of disturbed habitats, on Kure, Midway, and Pearl and Hermes atolls, French Frigate Shoals, Nihoa, Ka‘ula, Lehua, and documented from all of the main islands except Niihau. Widely naturalized prior to 1871 (Hillebr and, 1888).—Plate 35.