Gymnogramma pilosa Brack.; Coniogramme fraxinea sensu W. J. Rob., non (D. Don) Diels; Gymnogramma javanica (as Gymnogramme) sensu Hillebr., non Blume

Common names: lo'ulu

Endemic

Greek pilos, hair, felt, referring to the hairs on the upper surface of the pinnae.

Plants medium-sized, terrestrial. Rhizomes creeping, slender, 5-9 mm thick. Fronds 30-90 cm long. Stipes long. Blades 1-to 2-pinnate (lower 1-3 pinnae again 1-pinnate with 1-3(-5) pairs of large pinnules), light olive green, frond tips the same as lateral pinnae; rachises straw-colored, glossy, glabrous. Pinnae 3-7 pairs, stalked, alternate, ovate-lanceolate, 15-23 x 3.6-5 cm, margins crenulate with fine, cartilaginous teeth, upper surfaces glabrous, lower surfaces, costae, and veins with many scattered, fine, white, pointed hairs. Veins free, long, curved, 0-to 1(-2) forked (occasionally 2 veins joining). Sori along all veins and branches long-linear, to 2.2 em long, extending from costae to 3/4-7/8 distance to margins.

Locally common in mesic to wet forests, 610-2,070 m, all major islands.

Coniogramme pilosa is characterized by its distinctive light green color; 1-pin· nate fronds except for the basal 1 or 2 pinnae that may have large pinnules; frond tips with the same shape as the lateral pinnae; and naked, long, linear sori on every vein of the fertile fronds.