Wagner, Warren L., Derral R. Herbst and S. H. Sohmer. 1999. Manual of the Flowering Plants of Hawai'i (Bernice Pauahi Bishop Museum Special Publication, 2 Vol. Set). Honolulu, HI
Flora of Hawaii:
Distribution Native to Asia and Africa, widely cultivated and now spread from the Atlantic coast of Africa throughout the Pacific and more recently to the Neotropics. In the Marquesas a Polynesian introduction, now naturalized. In the Hawaiian Islands, a Polynesian introduction on Kaua`i, O`ahu, Moloka`i, Maui, Hawai`i.
Habit Stems twining to the left, slightly angled, up to 30 m long, glabrous, never with prickles, tubers nearly always solitary, 5-8 cm in diameter.
Leaves Leaves simple, alternate, blades broadly ovate cordate, 4-20 cm long, usually 3-18 cm wide, 7-11 nerved, glabrous, petioles 6-16 cm long, axils usually with bulbils.
Flowers Staminate flowers widely spaced, in slender, simple or fasciculate spikes or elongate panicles, each flower subtended by 2 unequal, ovate bracts, tepals white, lanceolate, 2.5-3 mm long, stamens 6, all fertile, ovary rudimentary. Pistillate flowers in pendent, rather stiff, simple spikes, these 3-6 in fascicles, tepals white, yellow with age, staminodes 6, stigmas irregularly 2-lobed.
Fruit Capsules ca. 25 mm long, with wings ca. 8 mm wide.
Seeds Seeds winged on basal side only.
Chromosomes 2n = 36, 40, 54, 60, 70, 80, 98 100.
Notes Naturalized and propagating vegetatively by means of stem tubers that fall to the ground and grow readily.