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  Animalia
      Arthropoda
        Hexapoda
            Insecta
              Pterygota
                Neoptera
                    Coleoptera
                      Polyphaga
                        Scarabaeiformia
                          Scarabaeoidea
                            Trogidae
                                     Cretomorgus
                                     Glyptotrox
                                             Glyptotrox boucomonti
                                             Glyptotrox brahminus
                                             Glyptotrox cambeforti
                                             Glyptotrox cambodjanus
                                             Glyptotrox dhaulagiri
                                             Glyptotrox doiinthanonensis
                                             Glyptotrox formosanus
                                             Glyptotrox foveicollis
                                             Glyptotrox frontera
                                             Glyptotrox hamatus
                                             Glyptotrox ineptus
                                             Glyptotrox insularis
                                             Glyptotrox kerleyi
                                             Glyptotrox kiuchii
                                             Glyptotrox mandli
                                             Glyptotrox matsudai
                                             Glyptotrox mutsuensis
                                             Glyptotrox opacotuberculatus
                                             Glyptotrox parvisetosus
                                             Glyptotrox poringensis
                                             Glyptotrox simi
                                             Glyptotrox spinulosus
                                             Glyptotrox sugayai
                                             Glyptotrox taiwanus
                                             Glyptotrox terrestris
                                             Glyptotrox tibialis
                                             Glyptotrox uenoi
                                             Glyptotrox yamayai
                                             Glyptotrox yangi
                                     Omorgus
                                             Omorgus acinus
                                             Omorgus alatus
                                             Omorgus alius
                                             Omorgus alternans
                                             Omorgus amictus
                                             Omorgus amitinus
                                             Omorgus aphanocephalus
                                             Omorgus asper
                                             Omorgus asperulatus
                                             Omorgus augustae
                                             Omorgus australasiae
                                             Omorgus baccatus
                                             Omorgus bachorum
                                             Omorgus badeni
                                             Omorgus batesi
                                             Omorgus bayoni
                                             Omorgus benadirensis
                                             Omorgus birmanicus
                                             Omorgus borgognoi
                                             Omorgus borrei
                                             Omorgus brucki
                                             Omorgus candezei
                                             Omorgus candidus
                                             Omorgus capillaceus
                                             Omorgus capillamentis
                                             Omorgus carinatus
                                             Omorgus carinicollis
                                             Omorgus chinensis
                                             Omorgus ciliatus
                                             Omorgus consanguineus
                                             Omorgus costatus
                                             Omorgus crotchi
                                             Omorgus curvipes
                                             Omorgus demarzi
                                             Omorgus denticulatus
                                             Omorgus dilaticollis
                                             Omorgus discedens
                                             Omorgus dohrni
                                             Omorgus drumonti
                                             Omorgus elderi
                                             Omorgus elevatus
                                             Omorgus elongatus
                                             Omorgus endroedyi
                                             Omorgus euclensis
                                             Omorgus expansus
                                             Omorgus eyrensis
                                             Omorgus fenestrellus
                                             Omorgus foveolatus
                                             Omorgus frater
                                             Omorgus freyi
                                             Omorgus fuliginosus
                                             Omorgus funestus
                                             Omorgus gemmatus
                                             Omorgus genieri
                                             Omorgus gigas
                                             Omorgus granulatus
                                             Omorgus granuliceps
                                             Omorgus guttalis
                                             Omorgus haagi
                                             Omorgus howdenorum
                                             Omorgus howelli
                                             Omorgus inclusus
                                             Omorgus indicus
                                             Omorgus indigenus
                                             Omorgus inermis
                                             Omorgus inflatus
                                             Omorgus insignicollis
                                             Omorgus insignis
                                             Omorgus italicus
                                             Omorgus lindemannae
                                             Omorgus lobicollis
                                             Omorgus loxus
                                             Omorgus lucidus
                                             Omorgus lugubris
                                             Omorgus maindroni
                                             Omorgus maissouri
                                             Omorgus mariae
                                             Omorgus mariettae
                                             Omorgus marshalli
                                             Omorgus melancholicus
                                             Omorgus mentitor
                                             Omorgus mictlensis
                                             Omorgus mollis
                                             Omorgus monachus
                                             Omorgus monteithi
                                             Omorgus mutabilis
                                             Omorgus nanningensis
                                             Omorgus nigroscobinus
                                             Omorgus niloticus
                                               Omorgus niloticus subsp. desertorum
                                             Omorgus nocheles
                                             Omorgus nodicollis
                                             Omorgus nodosus
                                             Omorgus obesus
                                             Omorgus omacanthus
                                             Omorgus ovalis
                                             Omorgus pampeanus
                                             Omorgus parvicollis
                                             Omorgus pastillarius
                                             Omorgus pauliani
                                             Omorgus pellosomus
                                             Omorgus perhispidus
                                             Omorgus persuberosus
                                             Omorgus ponderosus
                                             Omorgus principalis
                                             Omorgus procerus
                                             Omorgus punctatus
                                             Omorgus quadridens
                                             Omorgus quadrinodosus
                                             Omorgus radula
                                             Omorgus regalis
                                             Omorgus reiterorum
                                             Omorgus rimulosus
                                             Omorgus rodriguezae
                                             Omorgus rotundulus
                                             Omorgus rubricans
                                             Omorgus rusticus
                                             Omorgus salebrosus
                                             Omorgus satorui
                                             Omorgus scabrosus
                                             Omorgus scutellaris
                                             Omorgus semicostatus
                                             Omorgus senegalensis
                                             Omorgus setosipennis
                                             Omorgus spatulatus
                                             Omorgus squalidus
                                             Omorgus squamosus
                                             Omorgus strzeleckensis
                                             Omorgus subcarinatus
                                             Omorgus suberosus
                                             Omorgus tasmanicus
                                             Omorgus tatei
                                             Omorgus tessellatus
                                             Omorgus testudo
                                             Omorgus texanus
                                             Omorgus tomentosus
                                             Omorgus trilobus
                                             Omorgus tuberosus
                                             Omorgus tytus
                                             Omorgus umbonatus
                                             Omorgus undaraensis
                                             Omorgus unguicularis
                                             Omorgus varicosus
                                             Omorgus verrucosus
                                             Omorgus villosus
                                             Omorgus vladislavi
                                             Omorgus wittei
                                             Omorgus zumpti
                                     Paratrox
                                     Phoberus
                                             Phoberus aculeatus
                                             Phoberus arcuatus
                                             Phoberus braacki
                                             Phoberus brincki
                                             Phoberus caffer
                                               Phoberus caffer subsp. lilianae
                                             Phoberus capensis
                                             Phoberus consimilis
                                             Phoberus cyrtus
                                             Phoberus disjunctus
                                             Phoberus elmariae
                                             Phoberus gunki
                                             Phoberus herminae
                                             Phoberus horridus
                                             Phoberus levis
                                             Phoberus luridus
                                             Phoberus miliarius
                                             Phoberus montanus
                                             Phoberus mozalae
                                             Phoberus nama
                                             Phoberus nanniscus
                                             Phoberus nasutus
                                             Phoberus natalensis
                                             Phoberus necopinus
                                             Phoberus ngomensis
                                             Phoberus nigrociliatus
                                               Phoberus nigrociliatus subsp. nyansanus
                                               Phoberus nigrociliatus subsp. nyassicus
                                             Phoberus nyansanus
                                             Phoberus nyassicus
                                             Phoberus nyikanus
                                             Phoberus penicillatus
                                             Phoberus perrieri
                                             Phoberus planicollis
                                             Phoberus puncticollis
                                             Phoberus pusillus
                                             Phoberus quadricostatus
                                             Phoberus ranotsaraensis
                                             Phoberus rhyparoides
                                             Phoberus rudebecki
                                             Phoberus squamiger
                                             Phoberus sternbergi
                                             Phoberus strigosus
                                             Phoberus sulcatus
                                             Phoberus talpa
                                             Phoberus youngai
                                     Polynoncus
                                             Polynoncus aeger
                                             Polynoncus aricensis
                                             Polynoncus bifurcatus
                                             Polynoncus brasiliensis
                                             Polynoncus brevicollis
                                             Polynoncus bullatus
                                             Polynoncus burmeisteri
                                             Polynoncus chilensis
                                             Polynoncus diffluens
                                             Polynoncus ecuadorensis
                                             Polynoncus erugatus
                                             Polynoncus galapagoensis
                                             Polynoncus gemmifer
                                             Polynoncus gemmingeri
                                             Polynoncus gibberosus
                                             Polynoncus gordoni
                                             Polynoncus guttifer
                                             Polynoncus haafi
                                             Polynoncus hemisphaericus
                                             Polynoncus juglans
                                             Polynoncus longitarsis
                                             Polynoncus mirabilis
                                             Polynoncus neuquen
                                             Polynoncus parafurcatus
                                             Polynoncus patagonicus
                                             Polynoncus patriciae
                                             Polynoncus pedestris
                                             Polynoncus peruanus
                                             Polynoncus pilularius
                                             Polynoncus sallei
                                             Polynoncus seymourensis
                                             Polynoncus tenebrosus
                                     Prototrox
                                     Trox
                                             Trox abei
                                             Trox acanthinus
                                             Trox aequalis
                                             Trox affinis
                                             Trox alpigenus
                                             Trox aproximans
                                             Trox asper
                                             Trox atrox
                                             Trox boucomonti
                                             Trox brahminus
                                             Trox cadaverinus
                                               Trox cadaverinus subsp. komareki
                                             Trox cambeforti
                                             Trox cambodjanus
                                             Trox capensis
                                             Trox capillaris
                                             Trox carinatus
                                             Trox clathratus
                                             Trox confluens
                                             Trox contractus
                                             Trox coracinus
                                             Trox cotodognanensis
                                             Trox cribrum
                                             Trox cyrenaicus
                                             Trox dhaulagiri
                                             Trox doiinthanonensis
                                             Trox elkantaraensis
                                             Trox eversmanni
                                             Trox eximius
                                             Trox fabricii
                                             Trox fascicularis
                                               Trox fascicularis subsp. rowei
                                             Trox fascifer
                                             Trox floridanus
                                             Trox formosanus
                                             Trox foveicollis
                                             Trox frontera
                                             Trox fuliginosus
                                             Trox gansuensis
                                             Trox gemmulatus
                                             Trox granulipennis
                                             Trox hamata
                                             Trox hamatus
                                             Trox hispanicus
                                             Trox hispidus
                                               Trox hispidus subsp. niger
                                             Trox horiguchii
                                             Trox howelli
                                             Trox inadai
                                             Trox ineptus
                                             Trox inflatus
                                             Trox insularis
                                             Trox iranicus
                                             Trox jeanae
                                             Trox kerleyi
                                             Trox kiuchii
                                             Trox klapperichi
                                             Trox koreanus
                                             Trox kyotensis
                                             Trox lama
                                             Trox laticollis
                                             Trox leonardii
                                             Trox litoralis
                                             Trox lutosus
                                             Trox mandli
                                             Trox martini
                                             Trox matsudai
                                             Trox maurus
                                             Trox mitis
                                             Trox mixtus
                                             Trox monachus
                                             Trox morticinii
                                               Trox morticinii subsp. eximius
                                             Trox mutsuensis
                                             Trox niger
                                             Trox niisatorui
                                             Trox niponensis
                                             Trox nodosus
                                             Trox nodulosus
                                             Trox nohirai
                                             Trox opacotuberculatus
                                             Trox parvisetosus
                                             Trox paulseni
                                             Trox perlatus
                                             Trox perrisii
                                             Trox placosalinus
                                             Trox plicatus
                                             Trox poggii
                                             Trox poringensis
                                             Trox punctatus
                                             Trox quadrimaculatus
                                             Trox robinsoni
                                             Trox rubricans
                                             Trox sabulosus
                                               Trox sabulosus subsp. fujiokai
                                             Trox scaber
                                             Trox scabra
                                             Trox scabrosus
                                             Trox scutellaris
                                             Trox serrulatus
                                             Trox setifer
                                             Trox sonorae
                                             Trox sordidatus
                                             Trox sordidus
                                             Trox spinulosus
                                               Trox spinulosus dentibius
                                               Trox spinulosus subsp. simi
                                             Trox stellatus
                                             Trox strandi
                                             Trox striatus
                                             Trox suberosus
                                             Trox sugayai
                                             Trox taiwanus
                                             Trox terrestris
                                             Trox tessellatus
                                             Trox texanus
                                             Trox tibialis
                                             Trox torpidus
                                             Trox transversus
                                             Trox tuberculatus
                                             Trox tytus
                                             Trox uenoi
                                               Trox uenoi subsp. matsumurai
                                             Trox umbonatus
                                             Trox unistriatus
                                             Trox ussuriensis
                                             Trox variolatus
                                             Trox vimmeri
                                             Trox yamayai
                                             Trox yangi
                                             Trox zoufali