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  Animalia

      Arthropoda

        Hexapoda

            Insecta

              Pterygota

                Neoptera

                    Coleoptera

                      Polyphaga

                        Scarabaeiformia

                          Scarabaeoidea

                            Dynastidae

                                     Aceratus

                                             Aceratus davus

                                             Aceratus phyllis

                                     Acrobolbia

                                             Acrobolbia macrophylla

                                     Actinobolus

                                             Actinobolus alvarengai

                                             Actinobolus barbatus

                                             Actinobolus degallieri

                                             Actinobolus emarginatus

                                             Actinobolus pilosus

                                             Actinobolus radians

                                             Actinobolus spiniformis

                                             Actinobolus trilobus

                                     Adelaeus

                                             Adelaeus anachoretus

                                     Adoryphorus

                                             Adoryphorus canei

                                             Adoryphorus couloni

                                             Adoryphorus mellori

                                     Aegopsis

                                             Aegopsis bolboceridus

                                             Aegopsis chaminadei

                                             Aegopsis curvicornis

                                             Aegopsis peruvianus

                                     Agaocephala

                                             Agaocephala bicuspis

                                             Agaocephala brasiliana

                                             Agaocephala cornigera

                                             Agaocephala duponti

                                             Agaocephala inermicollis

                                             Agaocephala mannerheimi

                                             Agaocephala margaridae

                                             Agaocephala melolonthida

                                             Agaocephala urus

                                     Alissonotum

                                             Alissonotum africanum

                                             Alissonotum amploides

                                             Alissonotum binodulum

                                             Alissonotum camerunum

                                             Alissonotum consonense

                                             Alissonotum crassum

                                             Alissonotum cribratellum

                                             Alissonotum elongatum

                                             Alissonotum impressicolle

                                             Alissonotum lerui

                                             Alissonotum nitens

                                             Alissonotum parvum

                                             Alissonotum pauper

                                             Alissonotum pauperum

                                             Alissonotum piceum

                                               Alissonotum piceum subsp. besucheti

                                               Alissonotum piceum subsp. fujiokai

                                             Alissonotum pittinoi

                                             Alissonotum quadridens

                                             Alissonotum rangunense

                                             Alissonotum simile

                                             Alissonotum suzukii

                                             Alissonotum tibetense

                                             Alissonotum yamayai

                                             Alissonotum yunnaense

                                     Allomyrina

                                             Allomyrina davidis

                                             Allomyrina dichotoma

                                             Allomyrina pfeifferi

                                               Allomyrina pfeifferi subsp. celebensis

                                     Allophileurinus

                                             Allophileurinus cavifrons

                                             Allophileurinus mediopunctatus

                                     Amblyoproctus

                                             Amblyoproctus amazonicus

                                             Amblyoproctus arianarum

                                             Amblyoproctus bituberculatus

                                             Amblyoproctus boondocksius

                                             Amblyoproctus centroamericanus

                                             Amblyoproctus chalumeaui

                                             Amblyoproctus cornutus

                                             Amblyoproctus dechambrei

                                             Amblyoproctus ocellatus

                                             Amblyoproctus piliger

                                             Amblyoproctus rugosus

                                             Amblyoproctus squamosus

                                             Amblyoproctus torulosus

                                     Ampotis

                                             Ampotis alticola

                                     Anomacaulus

                                             Anomacaulus fulvovestitus

                                     Anomalomorpha

                                             Anomalomorpha anthracina

                                             Anomalomorpha giveni

                                             Anomalomorpha monteithi

                                     Anoronotum

                                             Anoronotum rufum

                                     Antodon

                                             Antodon goryi

                                     Aphonides

                                             Aphonides turbator

                                     Aphonodelus

                                             Aphonodelus coriaceipennis

                                             Aphonodelus mraci

                                     Archophanes

                                             Archophanes cratericollis

                                     Arriguttia

                                             Arriguttia bolivari

                                             Arriguttia brevissima

                                     Augoderia

                                             Augoderia boliviana

                                             Augoderia freyi

                                             Augoderia ginglarisi

                                             Augoderia nitidula

                                               Augoderia nitidula subsp. yungana

                                     Augosoma

                                             Augosoma centaurus

                                             Augosoma hippocrates

                                     Australocus

                                             Australocus bicornutus

                                     Barutus

                                             Barutus hartmanni

                                     Beckius

                                             Beckius beccarii

                                               Beckius beccarii subsp. ryusuii

                                     Blabephorus

                                             Blabephorus pinguis

                                     Brachysiderus

                                             Brachysiderus benjamini

                                             Brachysiderus bertolossiorum

                                             Brachysiderus elyanae

                                             Brachysiderus goyanus

                                             Brachysiderus martinae

                                             Brachysiderus matogrossensis

                                             Brachysiderus mielkeorum

                                             Brachysiderus minicola

                                             Brachysiderus paranensis

                                               Brachysiderus paranensis subsp. lenorae

                                             Brachysiderus quadrimaculatus

                                               Brachysiderus quadrimaculatus subsp. tridentiger

                                     Calcitoryctes

                                     Calicnemis

                                             Calicnemis latreillei

                                             Calicnemis obesa

                                               Calicnemis obesa subsp. sardiniensis

                                             Calicnemis truncatifrons

                                     Callistemonus

                                             Callistemonus intrusus

                                     Calypsoryctes

                                             Calypsoryctes dynastoides

                                     Carneiola

                                             Carneiola zealandica

                                     Carneodon

                                             Carneodon bidens

                                             Carneodon glauerti

                                             Carneodon intermedius

                                             Carneodon laevicollis

                                             Carneodon laevipennis

                                             Carneodon laevis

                                             Carneodon meyricki

                                             Carneodon occidentalis

                                             Carneodon pecuarius

                                             Carneodon simplex

                                     Carneoryctes

                                             Carneoryctes ater

                                             Carneoryctes brittoni

                                             Carneoryctes griseopilosus

                                             Carneoryctes minchami

                                             Carneoryctes monstrosus

                                             Carneoryctes nigripennis

                                             Carneoryctes peterseni

                                             Carneoryctes pimbus

                                             Carneoryctes psilus

                                             Carneoryctes semicalvus

                                             Carneoryctes sulcatus

                                             Carneoryctes tectus

                                             Carneoryctes tricornutus

                                             Carneoryctes trifidus

                                             Carneoryctes truncatus

                                             Carneoryctes wingarus

                                     Cavonus

                                             Cavonus acutifrons

                                             Cavonus armatus

                                             Cavonus bidens

                                             Cavonus leai

                                             Cavonus niger

                                             Cavonus sculpturatus

                                             Cavonus sharpi

                                     Caymania

                                             Caymania nitidissima

                                     Ceratophileurus

                                             Ceratophileurus lemoulti

                                     Ceratoryctoderus

                                             Ceratoryctoderus armatus

                                             Ceratoryctoderus borchmanni

                                             Ceratoryctoderus candezei

                                             Ceratoryctoderus mototanii

                                     Chalcasthenes

                                             Chalcasthenes divinus

                                             Chalcasthenes pulcher

                                             Chalcasthenes squamigera

                                             Chalcasthenes styracoceros

                                     Chalcocrates

                                             Chalcocrates bacchusi

                                             Chalcocrates borchmanni

                                             Chalcocrates bretoni

                                             Chalcocrates felschei

                                             Chalcocrates uniformis

                                     Chalepides

                                             Chalepides alliaceus

                                             Chalepides anomalus

                                             Chalepides barbatus

                                             Chalepides carinatus

                                             Chalepides comes

                                             Chalepides dilatatus

                                             Chalepides eucephalus

                                             Chalepides euhirtus

                                             Chalepides fuliginosus

                                             Chalepides howdenorum

                                             Chalepides hydrophiloides

                                             Chalepides luridus

                                             Chalepides narcisoi

                                             Chalepides osunai

                                             Chalepides unicolor

                                     Cheiroplatys

                                             Cheiroplatys amphioxus

                                             Cheiroplatys bifossus

                                             Cheiroplatys borealis

                                             Cheiroplatys carnei

                                             Cheiroplatys castaneus

                                             Cheiroplatys excavatus

                                             Cheiroplatys inconspicuus

                                             Cheiroplatys laevicollis

                                             Cheiroplatys latipes

                                             Cheiroplatys nitidus

                                             Cheiroplatys solidus

                                             Cheiroplatys volsellus

                                     Chiliphileurus

                                             Chiliphileurus tuberculatus

                                     Clyster

                                             Clyster hirokiharukae

                                             Clyster itys

                                             Clyster rectus

                                             Clyster retusus

                                               Clyster retusus subsp. nakanei

                                             Clyster rosaemariae

                                             Clyster scaurus

                                             Clyster trachypygus

                                             Clyster trinodosus

                                     Cnemidophileurus

                                             Cnemidophileurus personatus

                                     Coelosis

                                             Coelosis bicornis

                                             Coelosis biloba

                                             Coelosis bourgini

                                             Coelosis denticornis

                                             Coelosis hippocrates

                                             Coelosis inermis

                                             Coelosis sylvanus

                                     Coenoryctoderus

                                             Coenoryctoderus candezei

                                             Coenoryctoderus robustus

                                     Colacus

                                             Colacus bicolor

                                             Colacus morio

                                             Colacus moroni

                                     Collagenus

                                             Collagenus dasysternus

                                     Coptognathus

                                             Coptognathus attila

                                             Coptognathus crassipes

                                     Corynophyllus

                                             Corynophyllus andersoni

                                             Corynophyllus antennalis

                                             Corynophyllus arrowi

                                             Corynophyllus curvicornis

                                             Corynophyllus debilis

                                             Corynophyllus deserti

                                             Corynophyllus dorrigoensis

                                             Corynophyllus elricus

                                             Corynophyllus fortnumi

                                             Corynophyllus interocularis

                                             Corynophyllus macdougalli

                                             Corynophyllus major

                                             Corynophyllus modestus

                                             Corynophyllus pygidialis

                                             Corynophyllus teinogenoides

                                             Corynophyllus tulanus

                                     Cryptodus

                                             Cryptodus bilobus

                                             Cryptodus caviceps

                                             Cryptodus concentricus

                                             Cryptodus costulipennis

                                             Cryptodus cycnorum

                                             Cryptodus debilis

                                             Cryptodus foveatus

                                             Cryptodus fraternus

                                             Cryptodus gigas

                                             Cryptodus goerlingi

                                             Cryptodus grossipes

                                             Cryptodus matthewsi

                                             Cryptodus oblongoporus

                                             Cryptodus obscurus

                                             Cryptodus olivieri

                                             Cryptodus papuanus

                                             Cryptodus paradoxus

                                             Cryptodus passaloides

                                             Cryptodus piceus

                                             Cryptodus protensus

                                             Cryptodus relictus

                                             Cryptodus rotundicollis

                                             Cryptodus tasmannianus

                                             Cryptodus variolosus

                                     Cyphonistes

                                             Cyphonistes arrowi

                                             Cyphonistes brachti

                                             Cyphonistes burmeisteri

                                             Cyphonistes camurus

                                               Cyphonistes camurus subsp. bouyeri

                                             Cyphonistes corniculatus

                                               Cyphonistes corniculatus subsp. laevis

                                               Cyphonistes corniculatus subsp. transvaalensis

                                             Cyphonistes gasanus

                                             Cyphonistes glabricollis

                                             Cyphonistes impressicollis

                                             Cyphonistes inermicollis

                                             Cyphonistes ituriensis

                                             Cyphonistes latiusculus

                                             Cyphonistes letiranti

                                             Cyphonistes luluensis

                                             Cyphonistes roeri

                                             Cyphonistes sublaevis

                                             Cyphonistes tuberculifrons

                                             Cyphonistes vallatus

                                               Cyphonistes vallatus subsp. burgeoni

                                               Cyphonistes vallatus subsp. deserti

                                               Cyphonistes vallatus subsp. guineensis

                                               Cyphonistes vallatus subsp. rothschildi

                                               Cyphonistes vallatus subsp. septentrionis

                                     Dalgopus

                                             Dalgopus aethiopicus

                                             Dalgopus major

                                     Dasygnathus

                                             Dasygnathus blattocomes

                                             Dasygnathus dejeani

                                             Dasygnathus globosus

                                             Dasygnathus trituberculatus

                                     Democrates

                                             Democrates burmeisteri

                                             Democrates croesus

                                             Democrates lepiouffi

                                     Denhezia

                                             Denhezia caucacola

                                     Dichodontus

                                             Dichodontus birmanicus

                                             Dichodontus coronatus

                                             Dichodontus crassus

                                             Dichodontus croesus

                                             Dichodontus denticornis

                                             Dichodontus deruptus

                                             Dichodontus fissicornis

                                             Dichodontus grandis

                                             Dichodontus intermedius

                                             Dichodontus obesus

                                             Dichodontus pulchellus

                                             Dichodontus punctipennis

                                     Diloboderus

                                             Diloboderus abderus

                                     Dinoryctes

                                             Dinoryctes deguidei

                                             Dinoryctes truncaticollis

                                     Dipelicus

                                             Dipelicus acutus

                                             Dipelicus alveolatus

                                             Dipelicus bicarinatus

                                             Dipelicus bidens

                                             Dipelicus borneensis

                                             Dipelicus bovilli

                                             Dipelicus cantator

                                             Dipelicus cantori

                                             Dipelicus centratus

                                             Dipelicus daedalus

                                             Dipelicus deiphobus

                                               Dipelicus deiphobus subsp. loi

                                               Dipelicus deiphobus subsp. robustus

                                             Dipelicus duplex

                                             Dipelicus fastigatoides

                                             Dipelicus fastigatus

                                             Dipelicus furcatus

                                             Dipelicus geryon

                                             Dipelicus hircus

                                             Dipelicus indicus

                                             Dipelicus integriceps

                                             Dipelicus lacordairei

                                             Dipelicus minusculus

                                             Dipelicus mitratus

                                             Dipelicus monotuberculatus

                                             Dipelicus montrouzieri

                                             Dipelicus nasutus

                                             Dipelicus optatus

                                             Dipelicus ornatus

                                             Dipelicus pseudofastigatus

                                             Dipelicus pyramidalis

                                             Dipelicus quadratifer

                                             Dipelicus ritsemae

                                             Dipelicus sinuosus

                                             Dipelicus stanjakli

                                             Dipelicus triangularis

                                             Dipelicus tridentifer

                                             Dipelicus tuberosus

                                             Dipelicus ugobosiai

                                     Ebolowanius

                                             Ebolowanius mirei

                                     Enarotadius

                                             Enarotadius gressitti

                                     Endebius

                                             Endebius florensis

                                               Endebius florensis subsp. tanimbar

                                               Endebius florensis subsp. wetarensis

                                     Endroedianibe

                                             Endroedianibe bozzolai

                                             Endroedianibe xerozoyphius

                                     Enracius

                                             Enracius longipes

                                     Eophileurus

                                           [Trionychus]

                                             Eophileurus akitai

                                             Eophileurus assamensis

                                             Eophileurus baliensis

                                             Eophileurus baolocensis

                                             Eophileurus birmanicus

                                             Eophileurus borneensis

                                             Eophileurus celebensis

                                             Eophileurus chinensis

                                               Eophileurus chinensis subsp. irregularis

                                               Eophileurus chinensis subsp. okinawanus

                                             Eophileurus cingalensis

                                               Eophileurus cingalensis subsp. decatenatus

                                             Eophileurus confinis

                                             Eophileurus convexus

                                             Eophileurus dechambrei

                                             Eophileurus dentatus

                                             Eophileurus felschei

                                             Eophileurus fenicheli

                                             Eophileurus forsteri

                                             Eophileurus gracilis

                                             Eophileurus grossepunctatus

                                             Eophileurus heyrovskyi

                                             Eophileurus himalayanus

                                             Eophileurus howdeni

                                             Eophileurus iwasei

                                             Eophileurus javanus

                                             Eophileurus kachinensis

                                             Eophileurus malayanus

                                             Eophileurus malyi

                                             Eophileurus mentawaicus

                                             Eophileurus montanus

                                             Eophileurus multidentatus

                                             Eophileurus nicobarensis

                                             Eophileurus niii

                                             Eophileurus nilgirensis

                                             Eophileurus oblongus

                                             Eophileurus pectoralis

                                             Eophileurus perforatus

                                             Eophileurus planatus

                                             Eophileurus platypterus

                                             Eophileurus quadrigeminatus

                                             Eophileurus ryuheii

                                             Eophileurus sondaicus

                                             Eophileurus spatulicornis

                                             Eophileurus spinosus

                                             Eophileurus sukkiti

                                             Eophileurus sumbaianus

                                             Eophileurus takakuwai

                                             Eophileurus tanimbaricus

                                             Eophileurus tenuiformis

                                             Eophileurus tetraspermexitus

                                             Eophileurus thailandensis

                                             Eophileurus variolipennis

                                     Epironastes

                                             Epironastes abruptus

                                             Epironastes limbatus

                                             Epironastes nigrisetosus

                                     Eremobothynus

                                             Eremobothynus bicuspis

                                             Eremobothynus cornutus

                                             Eremobothynus hirsutus

                                     Erioscelis

                                             Erioscelis columbica

                                             Erioscelis emarginata

                                             Erioscelis peruana

                                             Erioscelis proba

                                             Erioscelis sobrina

                                     Eucopidocaulus

                                             Eucopidocaulus biintrusus

                                             Eucopidocaulus platypygus

                                             Eucopidocaulus tridentipes

                                     Eupatorus

                                             Eupatorus birmanicus

                                             Eupatorus endoi

                                             Eupatorus gracillicornis

                                               Eupatorus gracillicornis subsp. davidgohi

                                               Eupatorus gracillicornis subsp. edai

                                               Eupatorus gracillicornis subsp. kimioi

                                             Eupatorus hardwickei

                                             Eupatorus hardwickii

                                             Eupatorus koletta

                                             Eupatorus siamensis

                                               Eupatorus siamensis subsp. hirokii

                                             Eupatorus sukkiti

                                     Eutyctus

                                             Eutyctus deserti

                                     Gibboryctes

                                             Gibboryctes bollei

                                             Gibboryctes gracilicornis

                                             Gibboryctes szelenyii

                                             Gibboryctes waldenfelsi

                                     Gillaspytes

                                             Gillaspytes janzeni

                                     Gnathogolofa

                                             Gnathogolofa bicolor

                                     Goniophileurus

                                             Goniophileurus explanatus

                                             Goniophileurus femoratus

                                     Gorditus

                                             Gorditus rotundus

                                     Haplophileurus

                                             Haplophileurus caudipennis

                                             Haplophileurus dechambrei

                                             Haplophileurus uninodis

                                     Haploscapanes

                                             Haploscapanes australicus

                                             Haploscapanes barbarossa

                                             Haploscapanes inermis

                                             Haploscapanes papuanus

                                     Haplosoma

                                             Haplosoma ordinatum

                                     Harposcelis

                                             Harposcelis paradoxus

                                     Hatamus

                                             Hatamus gressittii

                                             Hatamus inarmatus

                                             Hatamus spinosus

                                             Hatamus tarsalis

                                     Heteroconus

                                             Heteroconus excellens

                                             Heteroconus fairmairei

                                             Heteroconus major

                                             Heteroconus muticus

                                             Heteroconus nanus

                                             Heteroconus paradoxus

                                             Heteroconus pauliani

                                             Heteroconus peyrierasi

                                     Heteroglobus

                                             Heteroglobus obesus

                                     Heterogomphus

                                             Heterogomphus achilles

                                             Heterogomphus aequatorius

                                             Heterogomphus aidoneus

                                             Heterogomphus amphitryon

                                             Heterogomphus arrowi

                                             Heterogomphus bicuspis

                                             Heterogomphus binodosus

                                             Heterogomphus bispinosus

                                             Heterogomphus bourcieri

                                             Heterogomphus carayoni

                                             Heterogomphus castaneus

                                             Heterogomphus chevrolati

                                             Heterogomphus consanguineus

                                             Heterogomphus consors

                                             Heterogomphus coriaceus

                                             Heterogomphus cribricollis

                                             Heterogomphus curvicornis

                                             Heterogomphus dejeani

                                             Heterogomphus dilaticollis

                                             Heterogomphus flohri

                                             Heterogomphus gibbosus

                                             Heterogomphus hiekei

                                             Heterogomphus hirticollis

                                             Heterogomphus hirtus

                                             Heterogomphus hopei

                                             Heterogomphus inarmatus

                                             Heterogomphus incornutus

                                             Heterogomphus julus

                                             Heterogomphus laticollis

                                             Heterogomphus mirabilis

                                             Heterogomphus mniszechi

                                             Heterogomphus monotuberculatus

                                             Heterogomphus ochoai

                                             Heterogomphus olsoufieffi

                                             Heterogomphus onorei

                                             Heterogomphus orsilochus

                                             Heterogomphus pauson

                                             Heterogomphus peruanus

                                             Heterogomphus pilosus

                                             Heterogomphus porioni

                                             Heterogomphus rubripennis

                                             Heterogomphus rugicollis

                                             Heterogomphus rugosus

                                             Heterogomphus schoenherri

                                             Heterogomphus telamon

                                             Heterogomphus thoas

                                             Heterogomphus ulysses

                                     Heteroligus

                                           [Aphonoproctus]

                                             Heteroligus appius

                                             Heteroligus bianoensis

                                             Heteroligus brevis

                                             Heteroligus falvopilosus

                                             Heteroligus geotrupinus

                                             Heteroligus meles

                                             Heteroligus minor

                                             Heteroligus miser

                                             Heteroligus pentodontinus

                                             Heteroligus peringueyi

                                             Heteroligus planifrons

                                             Heteroligus rojkoffi

                                             Heteroligus sebastiani

                                             Heteroligus truncaticeps

                                             Heteroligus vagabundus

                                     Heterostriatus

                                             Heterostriatus lerui

                                     Hexodon

                                             Hexodon griseosericans

                                             Hexodon kochi

                                             Hexodon latissimum

                                             Hexodon minutum

                                             Hexodon montandoni

                                             Hexodon patella

                                             Hexodon quadriplagiatum

                                             Hexodon reticulatum

                                             Hexodon unicolor

                                               Hexodon unicolor subsp. convexus

                                               Hexodon unicolor subsp. gigas

                                               Hexodon unicolor subsp. megaspilota

                                             Hexodon unicostatum

                                               Hexodon unicostatum subsp. simplex

                                     Hiekeianus

                                             Hiekeianus halophilus

                                             Hiekeianus tuberculatus

                                     Hispanioryctes

                                             Hispanioryctes paulseni

                                             Hispanioryctes rugulopygus

                                             Hispanioryctes sculptilis

                                             Hispanioryctes wittmeri

                                     Homoeomorphus

                                             Homoeomorphus nitidus

                                             Homoeomorphus rufipes

                                     Homophileurus

                                             Homophileurus aequatorius

                                             Homophileurus amazonicus

                                             Homophileurus cubanus

                                             Homophileurus integer

                                             Homophileurus luedeckei

                                             Homophileurus luederwaldti

                                             Homophileurus planus

                                             Homophileurus quadrituberculatus

                                             Homophileurus tricuspis

                                             Homophileurus waldenfelsi

                                     Hoploryctoderus

                                             Hoploryctoderus chinensis

                                             Hoploryctoderus tridens

                                               Hoploryctoderus tridens subsp. caledonicus

                                               Hoploryctoderus tridens subsp. pinorum

                                     Horridocalia

                                             Horridocalia delislei

                                     Hovophileurus

                                             Hovophileurus hovanus

                                             Hovophileurus latus

                                             Hovophileurus punctatostriatus

                                             Hovophileurus strigipennis

                                             Hovophileurus sulcicollis

                                     Hyboschema

                                             Hyboschema contractum

                                     Hylobothynus

                                             Hylobothynus arnaudi

                                             Hylobothynus columbianus

                                             Hylobothynus marielae

                                             Hylobothynus obesus

                                             Hylobothynus pontis

                                     Hyphoryctes

                                             Hyphoryctes maculatus

                                     Indieraligus

                                             Indieraligus michelii

                                     Irazua

                                             Irazua dilicra

                                     Licnostrategus

                                             Licnostrategus endymion

                                     Lonchotus

                                             Lonchotus aldabrensis

                                             Lonchotus ambrensis

                                             Lonchotus astovensis

                                             Lonchotus crassus

                                             Lonchotus endroedii

                                             Lonchotus latus

                                             Lonchotus lentus

                                             Lonchotus minor

                                             Lonchotus mohelicus

                                             Lonchotus politus

                                             Lonchotus punctatissimus

                                             Lonchotus rotundus

                                             Lonchotus rugosicollis

                                             Lonchotus sicardi

                                     Lycomedes

                                             Lycomedes bubeniki

                                             Lycomedes buckleyi

                                             Lycomedes burmeisteri

                                             Lycomedes hirtipes

                                             Lycomedes lydiae

                                             Lycomedes ohausi

                                             Lycomedes ramosus

                                             Lycomedes reichei

                                             Lycomedes salazari

                                             Lycomedes velutipes

                                     Macrocyphonistes

                                             Macrocyphonistes kolbeanus

                                     Marronus

                                             Marronus borbonicus

                                     Megaceras

                                             Megaceras bolivianus

                                             Megaceras brevis

                                             Megaceras briansaltini

                                             Megaceras celatum

                                             Megaceras crassum

                                             Megaceras endroedii

                                             Megaceras hoplites

                                             Megaceras inexspectatum

                                             Megaceras inflatum

                                             Megaceras jason

                                               Megaceras jason subsp. stuebeli

                                             Megaceras laevipenne

                                             Megaceras morpheus

                                             Megaceras pauliani

                                             Megaceras philoctetes

                                             Megaceras porioni

                                             Megaceras quadraticollis

                                             Megaceras remus

                                             Megaceras septentrionis

                                               Megaceras septentrionis subsp. angusticollis

                                     Megaceropsis

                                             Megaceropsis lecourti

                                             Megaceropsis quadridentatus

                                     Melanhyphus

                                             Melanhyphus clypealis

                                             Melanhyphus nanus

                                             Melanhyphus platygenioides

                                     Mellissius

                                             Mellissius adumbratus

                                             Mellissius eudoxus

                                             Mellissius oryctoides

                                             Mellissius popei

                                     Metanastes

                                             Metanastes australis

                                             Metanastes bicornis

                                               Metanastes bicornis subsp. papua

                                             Metanastes irregularis

                                             Metanastes picipes

                                             Metanastes vulgivagus

                                     Metaphileurus

                                             Metaphileurus lacunosus

                                             Metaphileurus nitidicollis

                                     Microphileurus

                                             Microphileurus caviceps

                                             Microphileurus subulo

                                     Microryctes

                                             Microryctes apicalis

                                             Microryctes confinis

                                             Microryctes dechambrei

                                             Microryctes hainanensis

                                             Microryctes kanarensis

                                             Microryctes katagirii

                                             Microryctes monodon

                                             Microryctes silvestrei

                                     Mictophileurus

                                             Mictophileurus punctulatus

                                     Mitracephala

                                             Mitracephala humboldti

                                             Mitracephala lachaumei

                                     Moraguesia

                                             Moraguesia champenoisi

                                     Musurgus

                                             Musurgus sabulicola

                                             Musurgus stridens

                                     Neocnecus

                                             Neocnecus minor

                                             Neocnecus punctata

                                     Neocorynophyllus

                                             Neocorynophyllus aputaringus

                                     Neodasygnathus

                                             Neodasygnathus corynophylloides

                                             Neodasygnathus davidsoni

                                             Neodasygnathus juba

                                     Neohyphus

                                             Neohyphus celebesus

                                             Neohyphus kleinschmidti

                                             Neohyphus philippinus

                                             Neohyphus quadridentatus

                                             Neohyphus semivelutinus

                                             Neohyphus suzukii

                                     Neometanastes

                                             Neometanastes hidakai

                                     Neonastes

                                             Neonastes glabricollis

                                             Neonastes uptoni

                                     Neoryctes

                                             Neoryctes galapagoensis

                                             Neoryctes moreti

                                             Neoryctes similis

                                             Neoryctes williamsi

                                     Nephrodopus

                                             Nephrodopus enigma

                                     Nimbacola

                                             Nimbacola lamottei

                                     Novapus

                                             Novapus adelaidae

                                             Novapus bifidus

                                             Novapus carnei

                                             Novapus crassus

                                             Novapus laticollis

                                             Novapus macfarlandi

                                             Novapus nitidus

                                             Novapus obscurus

                                             Novapus parvus

                                             Novapus rugosicollis

                                             Novapus simplex

                                             Novapus undarus

                                     Onychionyx

                                             Onychionyx australicus

                                             Onychionyx minor

                                             Onychionyx moluccanus

                                             Onychionyx opacipennis

                                     Orthocavonus

                                             Orthocavonus occidentalis

                                     Oryctoantiquus

                                     Oryctoderinus

                                             Oryctoderinus walfordorum

                                     Oryctoderus

                                             Oryctoderus albertisi

                                             Oryctoderus coronatus

                                             Oryctoderus godeffroyi

                                               Oryctoderus godeffroyi subsp. anguliceps

                                               Oryctoderus godeffroyi subsp. hebridarum

                                             Oryctoderus gracilior

                                             Oryctoderus latitarsis

                                             Oryctoderus sengokui

                                     Oryctophileurus

                                             Oryctophileurus armicollis

                                             Oryctophileurus guerrai

                                             Oryctophileurus nasicornis

                                             Oryctophileurus varicosus

                                     Oxyligyrus

                                             Oxyligyrus acutus

                                             Oxyligyrus cayennensis

                                             Oxyligyrus champenoisi

                                             Oxyligyrus contractus

                                             Oxyligyrus larssoni

                                             Oxyligyrus nasutus

                                             Oxyligyrus peruanus

                                             Oxyligyrus politus

                                             Oxyligyrus rostratus

                                             Oxyligyrus spitzi

                                             Oxyligyrus zoilus

                                     Pachyoryctes

                                             Pachyoryctes elongatus

                                             Pachyoryctes solidus

                                     Palaeophileurus

                                             Palaeophileurus brasiliensis

                                             Palaeophileurus carbo

                                             Palaeophileurus demezi

                                             Palaeophileurus erebus

                                             Palaeophileurus fallax

                                             Palaeophileurus marcusoni

                                             Palaeophileurus panamensis

                                             Palaeophileurus proximus

                                             Palaeophileurus sclateri

                                     Papuana

                                             Papuana angusta

                                             Papuana angustus

                                             Papuana armicollis

                                             Papuana biroi

                                             Papuana cheesmanae

                                             Papuana cognata

                                             Papuana coronata

                                             Papuana fascicula

                                             Papuana fortepunctata

                                             Papuana hardyi

                                             Papuana huebneri

                                               Papuana huebneri subsp. fallax

                                             Papuana inermis

                                             Papuana japenensis

                                             Papuana lansbergei

                                               Papuana lansbergei subsp. badia

                                             Papuana matsumotoi

                                             Papuana miyakei

                                             Papuana oryctoides

                                             Papuana philippinica

                                             Papuana sedlaceki

                                             Papuana splendens

                                             Papuana sumbana

                                             Papuana sumbawana

                                             Papuana szentivanyi

                                             Papuana tibialis

                                             Papuana timorensis

                                             Papuana trinodosa

                                             Papuana uninodis

                                             Papuana woodlarkiana

                                               Papuana woodlarkiana subsp. laevipennis

                                               Papuana woodlarkiana subsp. tenuistriata

                                     Paranodon

                                             Paranodon coquereli

                                     Paraphileurus

                                             Paraphileurus impressus

                                             Paraphileurus venezuelensis

                                             Paraphileurus ventricosus

                                     Parapucaya

                                             Parapucaya amazonica

                                             Parapucaya nodicollis

                                     Pareteronychus

                                             Pareteronychus dyscinetoides

                                     Parisomorphus

                                             Parisomorphus bouvieri

                                             Parisomorphus descarpentriesi

                                             Parisomorphus ganglbaueri

                                             Parisomorphus hilaris

                                             Parisomorphus niger

                                             Parisomorphus viettei

                                     Paroryctoderus

                                             Paroryctoderus cornutus

                                     Peltonotus

                                             Peltonotus adelphosimilis

                                             Peltonotus animus

                                             Peltonotus brunnipennis

                                             Peltonotus cybele

                                             Peltonotus deltamentum

                                             Peltonotus favonius

                                             Peltonotus fujiokai

                                             Peltonotus gracilipodus

                                             Peltonotus karubei

                                             Peltonotus kyojinus

                                             Peltonotus malayensis

                                             Peltonotus morio

                                             Peltonotus mushiyaus

                                             Peltonotus nasutus

                                             Peltonotus nethis

                                             Peltonotus podocrassus

                                             Peltonotus pruinosus

                                             Peltonotus rubripennis

                                             Peltonotus silvanus

                                             Peltonotus similis

                                             Peltonotus sisyrus

                                             Peltonotus suehirogarus

                                             Peltonotus talangensis

                                             Peltonotus tigerus

                                             Peltonotus vittatus

                                     Pentodina

                                             Pentodina peruviana

                                     Pentodontoschema

                                             Pentodontoschema caper

                                             Pentodontoschema capicola

                                             Pentodontoschema deceptor

                                             Pentodontoschema diversum

                                             Pentodontoschema endroedyyoungai

                                             Pentodontoschema nireum

                                     Pericoptus

                                             Pericoptus frontalis

                                             Pericoptus nitidulus

                                             Pericoptus punctatus

                                             Pericoptus studipus

                                             Pericoptus truncatus

                                     Phileucourtus

                                             Phileucourtus bicornutus

                                     Philoscaptus

                                             Philoscaptus bonariensis

                                     Phylliocephala

                                             Phylliocephala interioris

                                             Phylliocephala nigrohirta

                                     Pimelopus

                                             Pimelopus dubius

                                               Pimelopus dubius subsp. occidentalis

                                             Pimelopus fischeri

                                             Pimelopus minor

                                             Pimelopus noctis

                                             Pimelopus nothus

                                             Pimelopus porcellus

                                               Pimelopus porcellus subsp. crassus

                                               Pimelopus porcellus subsp. decipiens

                                             Pimelopus similis

                                     Piscoperus

                                             Piscoperus paracanicola

                                     Planophileurus

                                             Planophileurus broccus

                                             Planophileurus planicollis

                                             Planophileurus ypsilon

                                     Podischnus

                                             Podischnus agenor

                                             Podischnus oberthueri

                                             Podischnus sexdentatus

                                     Prionoryctes

                                             Prionoryctes breviusculus

                                             Prionoryctes camerunus

                                             Prionoryctes capreolus

                                             Prionoryctes intermedius

                                             Prionoryctes laticollis

                                             Prionoryctes rufopiceus

                                     Prosphileurus

                                             Prosphileurus liberianus

                                     Pseudocavonus

                                             Pseudocavonus antennalis

                                     Pseudohomonyx

                                             Pseudohomonyx apoensis

                                             Pseudohomonyx belitungensis

                                             Pseudohomonyx borneensis

                                             Pseudohomonyx crassus

                                             Pseudohomonyx extradentatus

                                             Pseudohomonyx javanus

                                               Pseudohomonyx javanus subsp. sumatrensis

                                             Pseudohomonyx luzonicus

                                             Pseudohomonyx mindroensis

                                             Pseudohomonyx negros

                                             Pseudohomonyx palavanus

                                             Pseudohomonyx samarensis

                                             Pseudohomonyx similis

                                     Pseudoryctes

                                             Pseudoryctes bidentifrons

                                             Pseudoryctes dispar

                                             Pseudoryctes etunus

                                             Pseudoryctes müllerianus

                                             Pseudoryctes semicrudus

                                             Pseudoryctes storeyi

                                             Pseudoryctes turritus

                                             Pseudoryctes validus

                                             Pseudoryctes wilsoni

                                     Pseudosyrichthus

                                             Pseudosyrichthus clathratus

                                             Pseudosyrichthus gerstaeckeri

                                             Pseudosyrichthus senegalensis

                                               Pseudosyrichthus senegalensis subsp. variolosus

                                     Pucaya

                                             Pucaya castanea

                                             Pucaya pulchra

                                     Pycnoschema

                                             Pycnoschema andrei

                                             Pycnoschema antinorii

                                             Pycnoschema antiorii

                                             Pycnoschema armatum

                                             Pycnoschema brittoni

                                             Pycnoschema corydon

                                             Pycnoschema cuspidatum

                                             Pycnoschema dechambrei

                                             Pycnoschema desfontainei

                                               Pycnoschema desfontainei subsp. ivoriense

                                             Pycnoschema diversum

                                               Pycnoschema diversum subsp. cavifrons

                                             Pycnoschema fortipes

                                             Pycnoschema hassoni

                                             Pycnoschema kisoa

                                             Pycnoschema lacordairei

                                             Pycnoschema mandibularis

                                             Pycnoschema mashunum

                                             Pycnoschema mirei

                                             Pycnoschema mossambicum

                                             Pycnoschema operculatum

                                             Pycnoschema parvicornis

                                             Pycnoschema politum

                                             Pycnoschema popei

                                             Pycnoschema prostome

                                             Pycnoschema prostomis

                                             Pycnoschema rawlinsi

                                             Pycnoschema schoutedeni

                                             Pycnoschema scrofa

                                             Pycnoschema simplicicolle

                                             Pycnoschema stricticeps

                                             Pycnoschema subulatum

                                             Pycnoschema suillum

                                             Pycnoschema uhligi

                                             Pycnoschema villiersi

                                     Rhizoplatodes

                                             Rhizoplatodes castaneipennis

                                               Rhizoplatodes castaneipennis subsp. foveiceps

                                             Rhizoplatodes cherlonneixi

                                     Rhizoplatys

                                             Rhizoplatys arabicus

                                             Rhizoplatys auriculatus

                                                   [Trionychus bituberculatus]

                                             Rhizoplatys canui

                                             Rhizoplatys cribarius

                                             Rhizoplatys maintyi

                                             Rhizoplatys mucronatus

                                               Rhizoplatys mucronatus subsp. cedrici

                                               Rhizoplatys mucronatus subsp. trituberculatus

                                             Rhizoplatys simplicitarsis

                                     Ruteloryctes

                                             Ruteloryctes bis

                                             Ruteloryctes morio

                                     Saccharoscaptus

                                             Saccharoscaptus laminifer

                                     Scapanes

                                             Scapanes affinis

                                             Scapanes australis

                                               Scapanes australis subsp. brevicornis

                                               Scapanes australis subsp. grossepunctatus

                                               Scapanes australis subsp. salomenensis

                                     Semanopterus

                                             Semanopterus angustatus

                                             Semanopterus calabyi

                                             Semanopterus leai

                                             Semanopterus rectangulus

                                             Semanopterus solidus

                                             Semanopterus subcostatus

                                             Semanopterus tricostatus

                                     Siralus

                                             Siralus insularis

                                     Syrichthodontus

                                             Syrichthodontus cribratus

                                             Syrichthodontus lydenburgi

                                             Syrichthodontus spurius

                                     Syrichthomorphus

                                             Syrichthomorphus termitophilus

                                     Syrichthoschema

                                             Syrichthoschema burgeoni

                                             Syrichthoschema katanganum

                                             Syrichthoschema vingerhoedti

                                     Syrictes

                                             Syrictes curvicornis

                                             Syrictes dentatus

                                             Syrictes rugulosus

                                             Syrictes syrichtus

                                     Talautoclyster

                                             Talautoclyster marikoae

                                     Tehuacania

                                             Tehuacania howdeni

                                     Teinogenys

                                             Teinogenys annectens

                                             Teinogenys aurilegulus

                                             Teinogenys calvus

                                             Teinogenys carinaticeps

                                             Teinogenys deceptus

                                             Teinogenys demarzi

                                             Teinogenys dives

                                             Teinogenys inermicollis

                                             Teinogenys inermis

                                             Teinogenys jenkinsi

                                             Teinogenys laevis

                                             Teinogenys metallicola

                                             Teinogenys mooniensis

                                             Teinogenys nitidus

                                             Teinogenys paradoxus

                                             Teinogenys richardsae

                                     Thronistes

                                             Thronistes rouxi

                                     Trichogomphus

                                             Trichogomphus acuticollis

                                             Trichogomphus bronchus

                                             Trichogomphus dechambrei

                                             Trichogomphus excavatus

                                               Trichogomphus excavatus subsp. fairmairei

                                             Trichogomphus galeatus

                                             Trichogomphus kobayashii

                                             Trichogomphus lunicollis

                                               Trichogomphus lunicollis subsp. fujiokai

                                             Trichogomphus martabani

                                             Trichogomphus mongol

                                             Trichogomphus pistiloides

                                             Trichogomphus quadridentatus

                                             Trichogomphus robustus

                                             Trichogomphus rongi

                                             Trichogomphus simson

                                               Trichogomphus simson subsp. selangorensis

                                             Trichogomphus vicinus

                                             Trichogomphus zangi

                                     Trioplus

                                             Trioplus cylindricus

                                     Trissodon

                                             Trissodon australasiae

                                             Trissodon bidentatus

                                             Trissodon bryanti

                                             Trissodon curtus

                                             Trissodon glaber

                                             Trissodon howdenorum

                                             Trissodon kiwanus

                                             Trissodon laevigatus

                                             Trissodon laticollis

                                             Trissodon nasutus

                                             Trissodon setosus

                                             Trissodon subopacus

                                             Trissodon tibialis

                                             Trissodon tudor

                                     Trypoxylus

                                             Trypoxylus dichotomus

                                               Trypoxylus dichotomus subsp. septentrionalis

                                               Trypoxylus dichotomus subsp. shennongji

                                               Trypoxylus dichotomus subsp. takarai

                                               Trypoxylus dichotomus subsp. tsuchiyai

                                               Trypoxylus dichotomus subsp. tsunobosonis

                                               Trypoxylus dichotomus subsp. xizangensis

                                             Trypoxylus kanamorii

                                     Wernoryctes

                                             Wernoryctes minettii

                                             Wernoryctes werneri

                                     Xenodorus

                                             Xenodorus basquini

                                             Xenodorus janus

                                     Xylotrupes

                                             Xylotrupes australicus

                                               Xylotrupes australicus subsp. darwinia

                                             Xylotrupes beckeri

                                               Xylotrupes beckeri subsp. intermedius

                                             Xylotrupes carinulus

                                             Xylotrupes clinias

                                               Xylotrupes clinias subsp. buru

                                             Xylotrupes damarensis

                                             Xylotrupes faber

                                             Xylotrupes falcatus

                                             Xylotrupes gideon

                                               Xylotrupes gideon subsp. borneensis

                                               Xylotrupes gideon subsp. lakorensis

                                               Xylotrupes gideon subsp. sawuensis

                                               Xylotrupes gideon subsp. sondaicus

                                             Xylotrupes inarmatus

                                             Xylotrupes lorquini

                                               Xylotrupes lorquini subsp. zideki

                                             Xylotrupes lumawigi

                                             Xylotrupes macleayi

                                               Xylotrupes macleayi subsp. asperulus

                                               Xylotrupes macleayi subsp. szekessyi

                                             Xylotrupes meridionalis

                                             Xylotrupes mirabilis

                                             Xylotrupes mniszechi

                                               Xylotrupes mniszechi subsp. beckeroides

                                               Xylotrupes mniszechi subsp. hainaniana

                                             Xylotrupes oudomxayicus

                                             Xylotrupes pachycera

                                             Xylotrupes pauliani

                                               Xylotrupes pauliani subsp. dayakorum

                                             Xylotrupes philippinensis

                                               Xylotrupes philippinensis subsp. boudanti

                                               Xylotrupes philippinensis subsp. peregrinus

                                             Xylotrupes pubescens

                                               Xylotrupes pubescens subsp. baumeisteri

                                               Xylotrupes pubescens subsp. beaudeti

                                               Xylotrupes pubescens subsp. gracilis

                                               Xylotrupes pubescens subsp. sibuyanensis

                                             Xylotrupes rindaae

                                             Xylotrupes socrates

                                               Xylotrupes socrates subsp. bourgini

                                               Xylotrupes socrates subsp. nitidus

                                               Xylotrupes socrates subsp. siamensis

                                               Xylotrupes socrates subsp. tonkinensis

                                             Xylotrupes striatopunctatus

                                             Xylotrupes sumatrensis

                                               Xylotrupes sumatrensis subsp. tanahmelayu

                                             Xylotrupes tadoana

                                             Xylotrupes taprobanus

                                               Xylotrupes taprobanus subsp. ganesha

                                             Xylotrupes telemachos

                                             Xylotrupes ulysses

                                             Xylotrupes wiltrudae